Professor of Biology Emeritus
Office: 1364E Spieth Hall
Phone (951) 827-5929
Degree:
Ph.D., Columbia University, 1955
Current research concerns the developmental biology of the frog Lepidobatrachus laevis. Work has been done in collaboration with other researchers and has involved egg membrane studies as well as an attempt to create gonadal chimaeras between two species of frogs (Lepidobatrachus and Xenopus). Prior research concerned the physiological ecology of desert anurans. Studies were done on terrestrial ecology of various species and the physiological aspects of water loss.